2018年,天网系统与人类的战争使世界陷入了末世般的境况,反抗军精神领袖约翰·康纳(克里斯蒂安·贝尔 Christian Bale 饰)带队执行一次任务的过程中获得了战胜天网的新思路,而天网系统适时推出了新的目标排名,康纳位居次席,榜首上康纳此时尚年青的父亲凯尔·瑞斯赫 然在列。与此同时,一名遗忘了自身过去经历的强壮男子马库斯(萨姆·沃辛顿 Sam Worthington 饰)出现在洛杉矶的废墟中,他与反抗军战士瑞斯偶然结识,并组队前往反抗军总部,不料途中遭机械人袭击,瑞斯被俘。落单的马库斯经由反抗军女战士布莱尔引路,迎来了与康纳命中注定的相遇,但人与机械的对立让康纳不得不视对方为敌人……   瑞斯在天网中心生死未卜,同时反抗军高层不肯推迟强行进攻计划,留给康纳的时间不再充裕,他只能选择进行一场赌上性命与荣誉的潜入行动。In 2003, in the Longview State Correctional Facility, the criminal Marcus Wright is on death row, and is convinced by the cancerous Dr. Serena Kogan to donate his body to her research and he accepts. In 2018, after an unsuccessful attack to a Skynet facility, only John Connor survives, but he discovers that Skynet is developing the powerful new model T-800. Out of the blue, Marcus appears naked and with amnesia in the location. Marcus befriends the teenager Kyle Reese and the girl Star who help him to survive the lethal machines and they travel together in a Jeep. Meanwhile the resistance discovers a signal that might turn-off the machines and John offers to test it. When Kyle is captured by a machine and brought to the Skynet headquarters, Marcus decides to help the youngster and heads to Skynet; on the way, he saves Blair Williams who suggests to him that he should meet John Connor first. But Marcus steps on a mine and is submitted to surgery, when a secret about his origins is disclosed.
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